Department of Biology
With a breadth of research areas, Clark’s biology doctoral program provides opportunities and flexibility to pursue research that benefits our understanding of the world within a connected community of scholars. Our research areas include molecular and cell biology, and ecology and evolution.
The program’s intimate size allows graduate students to collaborate, build relationships with and receive mentoring from world-class faculty. Close ties across the sciences, including with those enrolled in our biochemistry and molecular biology and chemistry doctoral programs, exposes students to a range of ideas and research outside their interest areas. Graduate students are guaranteed tuition remission and teaching assistantships for five years.
No Student Experiences for this program yet
Study ecosystem management online at Penn State. Gain in-demand skills in quantitative and spatial data analysis, project management, and leadership. Apply now for 2025!
Transform location intelligence into insight to protect our planet. Earn a Penn State GIS certificate online, on your time. Gain new career-ready technical skills to enhance your career.
Develop your skills in ecological research, quantitative analysis, GIS, and leadership to become an effective advocate for environmental conservation, education and justice. Apply for partial scholarships!